Updated on 30 March 2010
Total shares in issue 54,242,850
Shareholders with holdings over 3% were as follows:
University of Sheffield 12,895,541 23.77%
Cardiff University 10,997,541 20.27%
Invesco 5,570,363 10.27%
AXA Framlington 3,716,370 6.85%
Legal & General 3,325,926 6.13%
David Baynes 1,333,332 2.46%
Stuart Gall 1,333,332 2.46%
Peter Grant 1,333,332 2.46%